This modification of the ME_Gabor model has Gabor filters with Gaussian envelopes that are rotated in space-time to align with the orientation of the tilted sinusoidal grating, as depicted by McLean and Palmer (1994; their Fig 1A). The rotation in X-T corresponds to a rotation in spatio-temporal frequency space that causes the SF-TF tuning to be inseparable. Inseparable tuning has been observed in some direction selective V1 neurons in macaque (Priebe et al. 2006), but rarely in cat (McLean and Palmer, 1994), and has been associated with velocity, as opposed to SF and TF, tuning.
The signal, which is non-negative, is then combined with an additive offset, scaled, and half-wave rectified, the latter having no effect unless the signal scaling or offset has created negative values). The result is used to drive a Poisson spiking mechanism. The spikes are time shifted to simulate a neurobiological latency. See the model (.moo) files for the parameters that govern the computations.