The visual stimulus is processed (convolved) by four linear Gabor filters (icons show x-t slices of 3D filters) to yield the signals fpe (filter preferred even), fpo (filter preferred odd), fae (anti-preferred even) and fao (anti-preferred odd). Each filter output is squared, and the signals are added for each quadrature pair to give the motion energy in the preferred direction, mep, and the anti-preferred direction, mea. These two oppositely tuned ME signals are subtracted to give the opponent motion energy, meopp.

The opponent signal (meopp) is then offset, scaled and half-wave rectified, and it is used to drive a Poisson spiking mechanism. The spikes are time shifted to simulate a neurobiological latency. See the model (.moo) files for the parameters that govern these computations.